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Tolwani oversees the Gene Targeting Resource Center and Transgenic Services Laboratory of the Comparative Bioscience Center (CBC). The CBC provides expertise in animal models, research design and methodology, generation and evaluation of genetically engineered mouse models, and other collaborative efforts. It also provides training to Rockefeller investigators.

Tolwani holds a D.V.M. in veterinary medicine from Auburn University. After two years in private practice in Nashville, Tennessee, he turned to research, completing a Ph.D. in cellular and molecular pathology and a postdoc fellowship in laboratory animal/comparative medicine at the University of Alabama. For his research, he developed a mouse model to study a common illness in humans associated with an enzyme deficiency in fat metabolism. Tolwani later earned an M.Sx. degree in management from the Stanford Graduate School of Business.

Tolwani joined Rockefeller in 2007. From 1994 to 2007, he served on the faculty of the Stanford University School of Medicine, where his laboratory focused on understanding the molecular mechanisms of brain plasticity.