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B.S. in zoology, 1974University of California, Davis
Ph.D. in biochemistry, 1981 California Institute of Technology
California Institute of Technology, 1981–1985
Assistant Professor, 1986–1990Associate Professor, 1991–1995 Professor, 1995–2000Washington University School of Medicine
Professor, 2000–Scientific and Executive Director, Center for the Study of Hepatitis C, 2000–2018The Rockefeller University